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Abstract We use new cosmic microwave background (CMB) primary temperature and polarization anisotropy measurements from the Atacama Cosmology Telescope (ACT) Data Release 6 (DR6) to test foundational assumptions of the standard cosmological model, ΛCDM, and set constraints on extensions to it. We derive constraints from the ACT DR6 power spectra alone, as well as in combination with legacy data from thePlanckmission. To break geometric degeneracies, we include ACT andPlanckCMB lensing data and baryon acoustic oscillation data from DESI Year-1. To test the dependence of our results on non-ACT data, we also explore combinations replacingPlanckwithWMAPand DESI with BOSS, and further add supernovae measurements from Pantheon+ for models that affect the late-time expansion history. We verify the near-scale-invariance (running of the spectral indexdn_s/dlnk= 0.0062 ± 0.0052) and adiabaticity of the primordial perturbations. Neutrino properties are consistent with Standard Model predictions: we find no evidence for new light, relativistic species that are free-streaming (N_eff= 2.86 ± 0.13, which combined with astrophysical measurements of primordial helium and deuterium abundances becomesN_eff= 2.89 ± 0.11), for non-zero neutrino masses (∑m_ν< 0.089 eV at 95% CL), or for neutrino self-interactions. We also find no evidence for self-interacting dark radiation (N_idr< 0.134), or for early-universe variation of fundamental constants, including the fine-structure constant (α_EM/α_EM,0= 1.0043 ± 0.0017) and the electron mass (m_e/m_e,0= 1.0063 ± 0.0056). Our data are consistent with standard big bang nucleosynthesis (we findY_p= 0.2312 ± 0.0092), theCOBE/FIRAS-inferred CMB temperature (we findT_CMB= 2.698 ± 0.016 K), a dark matter component that is collisionless and with only a small fraction allowed as axion-like particles, a cosmological constant (w= -0.986 ± 0.025), and the late-time growth rate predicted by general relativity (γ= 0.663 ± 0.052). We find no statistically significant preference for a departure from the baseline ΛCDM model. In fits to models invoking early dark energy, primordial magnetic fields, or an arbitrary modified recombination history, we findH₀= 69.9^+0.8_-1.5, 69.1 ± 0.5, or 69.6 ± 1.0 km/s/Mpc, respectively; using BOSS instead of DESI BAO data reduces the central values of these constraints by 1–1.5 km/s/Mpc while only slightly increasing the error bars. In general, models introduced to increase the Hubble constant or to decrease the amplitude of density fluctuations inferred from the primary CMB are not favored over ΛCDM by our data.